Klaus-Robert Müller is a German physicist and computer scientist, most noted for his work in Machine Learning and Brain-Computer Interfaces. Dieter Waloszek is the author of Morphology, Ontogeny and Phylogeny of the Phosphatocopina - Crustacea - from the Upper Cambrian Orsten of Sweden, published by Wiley.

• Introduction 3“Orsten”-type fossils 3History of phosphatocopine research 3Prior assumptions on the systematics of Phosphatocopina. 3Soft parts and new perspectives about the phytogeny4Scope and aims 4Material and methods 4Material 4General information 4Statistical data 5Stratigraphy 6Methods 7Isolation of specimens (not done for this work) 7Preparation for SEM 8Measurements 8Terminology 8Systematics 10Crustacea Brunnich, 1772 13Phylogenetic analysis 13Results 13Hesslandona Milller, 1964. 15Hesslandona abdominalis Hinz-Schallreuter, 1998 16Hesslandona reichi Hinz-Schallreuter, 1993 16Hesslandona unisulcata Milller, 1982. 16Hesslandona necopina Milller, 1964. 58Hesslandona kinnekullensis Muller, 1964 67Hesslandona trituberculata (Lochman 8t Hu, 1960) Rushton, 1978 71Hesslandona ventrospinata Grundel in Grundel 8c Buchholz, 1981 77Hesslandona suecica n. sp 85Hesslandona angustata n. sp 89Hesslandona curvispina n. sp 93Hesslandona toreborgensis n. sp 104Trapezilites Hinz-Schallreuter, 1993 106Trapezilites minimus (Kummerow, 1931) Hinz-Schallreuter, 1993 106Waldoria Griindel in Griindel 8c Buchholz, 1981 113Waldoria rotundata Griindel in Griindel 8c Buchholz, 1981. 115Veldotron Grundel in Griindel 8c Buchholz, 1981 121Veldotron bratteforsa (Muller, 1964) Hinz-Schallreuter, 1993 123Falites Muller, 1964 128Falites fala Muller, 1964 131Vestrogothia Muller, 1964 137Vestrogothia spinata Muller, 1964 139Discussion 145Comparative morphology' 145Phylogenetic analysis of Phosphatocopina 159General remarks 159Reconstructed phytogeny of Phosphatocopina 160Phosphatocopina ("Phosphatocopida sp." + Euphosphatocopina new name 160Phosphatocopida sp 165Euphosphatocopina new taxon 165Vestrogothia spinata 168NN1 (Falites fala + Hesslandonina) 169Falites fala 169Hesslandonina (Cyclotronidae + Hesslandonidae) 170Cyclotronidae (Veldotron bratteforsa + Cyclotron lapworthi) 171Veldotron bratteforsa 171Cyclotron lapworthi 172Waldoria rotundata 172Hesslandonidae (Trapezilites minimus + Hesslandona) 173Trapezilites minimus 173Hesslandona 174Hesslandona unisulcata 174Dorsospinata new taxon 174NN3 (Hesslandona kinnekullensis+NN4) 175Hesslandona kinnekullensis 175NN4 (Hesslandona toreborgensis n. sp., H. angustata n. sp. + H. suecica n. sp.) 177Hesslandona toreborgensis n. sp 177Hesslandona suecica n. sp 177Hesslandona angustata n. sp 178NN5 (Hesslandona ventrospinata + NN6) 178Hesslandona ventrospinata............... 179NN6 (Hesslandona trituberculata + NN7) 179Hesslandona trituberculata.................. .....179NN7 (Hesslandona necopina + H. curvispina) 179Hesslandona necopina 180Hesslandona curvispina n. sp 180Consequences of the proposed phytogeny of Phosphatocopina 180Crustacean phytogeny 182Ground pattern of Crustacea 182The Ground pattern of Labrophora Siveter, Waloszek 8e Williams (2003) 185Ground pattern of Phosphatocopina 190Ground pattern of Eucrustacea 191Conclusions 194Future investigations and prospects 194Acknowledgements 194References 195Appendix A - Data for the phylogenetic analysis 203PAUP settings 203List of all coded characters 203Character matrix 206Appendix B — Synonymy of Phosphatocopine taxa 209Possible Phosphatocopine taxa 209Possible Phosphatocopines, described in open nomenclature 237

Klaus-Robert Müller is a German physicist and computer scientist, most noted for his work in Machine Learning and Brain-Computer Interfaces. Dieter Waloszek is the author of Morphology, Ontogeny and Phylogeny of the Phosphatocopina - Crustacea - from the Upper Cambrian Orsten of Sweden, published by Wiley.

• Introduction 3“Orsten”-type fossils 3History of phosphatocopine research 3Prior assumptions on the systematics of Phosphatocopina. 3Soft parts and new perspectives about the phytogeny4Scope and aims 4Material and methods 4Material 4General information 4Statistical data 5Stratigraphy 6Methods 7Isolation of specimens (not done for this work) 7Preparation for SEM 8Measurements 8Terminology 8Systematics 10Crustacea Brunnich, 1772 13Phylogenetic analysis 13Results 13Hesslandona Milller, 1964. 15Hesslandona abdominalis Hinz-Schallreuter, 1998 16Hesslandona reichi Hinz-Schallreuter, 1993 16Hesslandona unisulcata Milller, 1982. 16Hesslandona necopina Milller, 1964. 58Hesslandona kinnekullensis Muller, 1964 67Hesslandona trituberculata (Lochman 8t Hu, 1960) Rushton, 1978 71Hesslandona ventrospinata Grundel in Grundel 8c Buchholz, 1981 77Hesslandona suecica n. sp 85Hesslandona angustata n. sp 89Hesslandona curvispina n. sp 93Hesslandona toreborgensis n. sp 104Trapezilites Hinz-Schallreuter, 1993 106Trapezilites minimus (Kummerow, 1931) Hinz-Schallreuter, 1993 106Waldoria Griindel in Griindel 8c Buchholz, 1981 113Waldoria rotundata Griindel in Griindel 8c Buchholz, 1981. 115Veldotron Grundel in Griindel 8c Buchholz, 1981 121Veldotron bratteforsa (Muller, 1964) Hinz-Schallreuter, 1993 123Falites Muller, 1964 128Falites fala Muller, 1964 131Vestrogothia Muller, 1964 137Vestrogothia spinata Muller, 1964 139Discussion 145Comparative morphology' 145Phylogenetic analysis of Phosphatocopina 159General remarks 159Reconstructed phytogeny of Phosphatocopina 160Phosphatocopina ("Phosphatocopida sp." + Euphosphatocopina new name 160Phosphatocopida sp 165Euphosphatocopina new taxon 165Vestrogothia spinata 168NN1 (Falites fala + Hesslandonina) 169Falites fala 169Hesslandonina (Cyclotronidae + Hesslandonidae) 170Cyclotronidae (Veldotron bratteforsa + Cyclotron lapworthi) 171Veldotron bratteforsa 171Cyclotron lapworthi 172Waldoria rotundata 172Hesslandonidae (Trapezilites minimus + Hesslandona) 173Trapezilites minimus 173Hesslandona 174Hesslandona unisulcata 174Dorsospinata new taxon 174NN3 (Hesslandona kinnekullensis+NN4) 175Hesslandona kinnekullensis 175NN4 (Hesslandona toreborgensis n. sp., H. angustata n. sp. + H. suecica n. sp.) 177Hesslandona toreborgensis n. sp 177Hesslandona suecica n. sp 177Hesslandona angustata n. sp 178NN5 (Hesslandona ventrospinata + NN6) 178Hesslandona ventrospinata............... 179NN6 (Hesslandona trituberculata + NN7) 179Hesslandona trituberculata.................. .....179NN7 (Hesslandona necopina + H. curvispina) 179Hesslandona necopina 180Hesslandona curvispina n. sp 180Consequences of the proposed phytogeny of Phosphatocopina 180Crustacean phytogeny 182Ground pattern of Crustacea 182The Ground pattern of Labrophora Siveter, Waloszek 8e Williams (2003) 185Ground pattern of Phosphatocopina 190Ground pattern of Eucrustacea 191Conclusions 194Future investigations and prospects 194Acknowledgements 194References 195Appendix A - Data for the phylogenetic analysis 203PAUP settings 203List of all coded characters 203Character matrix 206Appendix B — Synonymy of Phosphatocopine taxa 209Possible Phosphatocopine taxa 209Possible Phosphatocopines, described in open nomenclature 237

Klaus-Robert Müller is a German physicist and computer scientist, most noted for his work in Machine Learning and Brain-Computer Interfaces. Dieter Waloszek is the author of Morphology, Ontogeny and Phylogeny of the Phosphatocopina - Crustacea - from the Upper Cambrian Orsten of Sweden, published by Wiley.

• Introduction 3“Orsten”-type fossils 3History of phosphatocopine research 3Prior assumptions on the systematics of Phosphatocopina. 3Soft parts and new perspectives about the phytogeny4Scope and aims 4Material and methods 4Material 4General information 4Statistical data 5Stratigraphy 6Methods 7Isolation of specimens (not done for this work) 7Preparation for SEM 8Measurements 8Terminology 8Systematics 10Crustacea Brunnich, 1772 13Phylogenetic analysis 13Results 13Hesslandona Milller, 1964. 15Hesslandona abdominalis Hinz-Schallreuter, 1998 16Hesslandona reichi Hinz-Schallreuter, 1993 16Hesslandona unisulcata Milller, 1982. 16Hesslandona necopina Milller, 1964. 58Hesslandona kinnekullensis Muller, 1964 67Hesslandona trituberculata (Lochman 8t Hu, 1960) Rushton, 1978 71Hesslandona ventrospinata Grundel in Grundel 8c Buchholz, 1981 77Hesslandona suecica n. sp 85Hesslandona angustata n. sp 89Hesslandona curvispina n. sp 93Hesslandona toreborgensis n. sp 104Trapezilites Hinz-Schallreuter, 1993 106Trapezilites minimus (Kummerow, 1931) Hinz-Schallreuter, 1993 106Waldoria Griindel in Griindel 8c Buchholz, 1981 113Waldoria rotundata Griindel in Griindel 8c Buchholz, 1981. 115Veldotron Grundel in Griindel 8c Buchholz, 1981 121Veldotron bratteforsa (Muller, 1964) Hinz-Schallreuter, 1993 123Falites Muller, 1964 128Falites fala Muller, 1964 131Vestrogothia Muller, 1964 137Vestrogothia spinata Muller, 1964 139Discussion 145Comparative morphology' 145Phylogenetic analysis of Phosphatocopina 159General remarks 159Reconstructed phytogeny of Phosphatocopina 160Phosphatocopina ("Phosphatocopida sp." + Euphosphatocopina new name 160Phosphatocopida sp 165Euphosphatocopina new taxon 165Vestrogothia spinata 168NN1 (Falites fala + Hesslandonina) 169Falites fala 169Hesslandonina (Cyclotronidae + Hesslandonidae) 170Cyclotronidae (Veldotron bratteforsa + Cyclotron lapworthi) 171Veldotron bratteforsa 171Cyclotron lapworthi 172Waldoria rotundata 172Hesslandonidae (Trapezilites minimus + Hesslandona) 173Trapezilites minimus 173Hesslandona 174Hesslandona unisulcata 174Dorsospinata new taxon 174NN3 (Hesslandona kinnekullensis+NN4) 175Hesslandona kinnekullensis 175NN4 (Hesslandona toreborgensis n. sp., H. angustata n. sp. + H. suecica n. sp.) 177Hesslandona toreborgensis n. sp 177Hesslandona suecica n. sp 177Hesslandona angustata n. sp 178NN5 (Hesslandona ventrospinata + NN6) 178Hesslandona ventrospinata............... 179NN6 (Hesslandona trituberculata + NN7) 179Hesslandona trituberculata.................. .....179NN7 (Hesslandona necopina + H. curvispina) 179Hesslandona necopina 180Hesslandona curvispina n. sp 180Consequences of the proposed phytogeny of Phosphatocopina 180Crustacean phytogeny 182Ground pattern of Crustacea 182The Ground pattern of Labrophora Siveter, Waloszek 8e Williams (2003) 185Ground pattern of Phosphatocopina 190Ground pattern of Eucrustacea 191Conclusions 194Future investigations and prospects 194Acknowledgements 194References 195Appendix A - Data for the phylogenetic analysis 203PAUP settings 203List of all coded characters 203Character matrix 206Appendix B — Synonymy of Phosphatocopine taxa 209Possible Phosphatocopine taxa 209Possible Phosphatocopines, described in open nomenclature 237