Heinrich Walter – författare

Phytomass and Primary Production of the Various Vegetational Zones and of the Entire Biosphere The biosphere is that thin layer at the earth''s surface in which living organisms exist and biological cycling takes place. It includes the upper horizons of the soil in which plants root, the atmosphere near the ground, (insofar as organisms penetrate this space), and all the surface waters. More than 99% of the earth''s biomass is phytomass, to which we shall limit our discussion. Amounts of phytomass are distinctly related to vegeta­ tional zones. Because accurate determination of phytomass and primary production is difficult, only gross estimates have been available until recently. However, in 1970, Bazilevich et al. published (in Russian) more accurate calculations, based on the rapidly accumulating literature, for the various thermal zones and bioclimatic regions of the earth. These authors calculated mean phyto­ mass and mean annual primary production for the various regions as dry mass (in tons) per hectare. On the basis of measurements of the areas covered by the individual regions, excluding rivers, lakes, glaciers, and permanent snow, total phytomass and total annual primary production for the various regions were obtained (see table). The sum of these figures is the phytomass and annual production of the land surface of the earth. In addition, the table gives corresponding data for the waters of the earth. The values involved are potential i. e. , they are based on natural vegetation uninfluenced by man.

We shall limit our observations to the conditions in natural ecosystems, since it would be beyond the scope of this book to embark upon a consideration of secondary, man-made ecosystems. 2. Classification of the Geo-biosphere into Zonobiomes The biosphere is the thin layer of the earth's surface to which the phenomena connected with living matter are confined. On land, this comprises the lowest layer of the atmosphere permanently inhabited by living organisms and into which plants extend, as well as the root-containing portion of the lithosphere, which we term the soil. Living organisms are also found in all bodies of water, to the very depths of the oceans. In a watery medium, however, cycling of material is achieved by means other than those on land, and the organisms (plankton) are so different that aquatic ecosystems have to be dealt with separately. The biosphere is therefore subdivided into (a) the geo-biosphere comprising terrestrial ecosystems, and (b) the hydro-biosphere, comprising aquatic ecosystems, which is the field of hydrobiologists (oceanographers and limnologists) . Our studies are confined to the geo-biosphere (Walter 1976), which constitutes the habitat of man and is, therefore, of special interest. The prevailing climate, being the primary independent factor in the environment, can be used as a basis for further subdivision of the geo-biosphere since the formation of soil and type of vegetation are dependent upon it (see p. 3), and it has not yet been substantially influenced by man.

By the end of the last century, the only region in all of Europe where the natural vegetation remained untouched over large expanses was Eastern Europe. Information on the vegetation of this region however, remained mostly unavailable to western scientists although the ecological approach to botanical research originated in Eastern Europe. Heinrich Walter was born in Russia, studied Natural Sciences - with emphasis on botany - in Odessa and is familiar with all relevant Russian literature on ecology. This volume thus offers scientists worldwide a unique and comprehensive survey of all relevant research, including the mostly unknown literature published in Russian.

By the end of the last century, the only region in all of Europe where the natural vegetation remained untouched over large expanses was Eastern Europe. Information on the vegetation of this region however, remained mostly unavailable to western scientists although the ecological approach to botanical research originated in Eastern Europe. Heinrich Walter was born in Russia, studied Natural Sciences - with emphasis on botany - in Odessa and is familiar with all relevant Russian literature on ecology. This volume thus offers scientists worldwide a unique and comprehensive survey of all relevant research, including the mostly unknown literature published in Russian.

We shall limit our observations to the conditions in natural ecosystems, since it would be beyond the scope of this book to embark upon a consideration of secondary, man-made ecosystems. 2. Classification of the Geo-biosphere into Zonobiomes The biosphere is the thin layer of the earth''s surface to which the phenomena connected with living matter are confined. On land, this comprises the lowest layer of the atmosphere permanently inhabited by living organisms and into which plants extend, as well as the root-containing portion of the lithosphere, which we term the soil. Living organisms are also found in all bodies of water, to the very depths of the oceans. In a watery medium, however, cycling of material is achieved by means other than those on land, and the organisms (plankton) are so different that aquatic ecosystems have to be dealt with separately. The biosphere is therefore subdivided into (a) the geo-biosphere comprising terrestrial ecosystems, and (b) the hydro-biosphere, comprising aquatic ecosystems, which is the field of hydrobiologists (oceanographers and limnologists) . Our studies are confined to the geo-biosphere (Walter 1976), which constitutes the habitat of man and is, therefore, of special interest. The prevailing climate, being the primary independent factor in the environment, can be used as a basis for further subdivision of the geo-biosphere since the formation of soil and type of vegetation are dependent upon it (see p. 3), and it has not yet been substantially influenced by man.

Des Menschen Werk auf Erden kann vergehen, Doch Leben stets im Wandel wird bestehen. Heinrich Walter The importance of ecology for the fate of mankind is receiving ever wider recognition. A syncretic-holistic approach to ecology was recently given unexpected support by the well-known atomic physicist and pupil of Heisenberg, Fritjov Capra. In his book The Turning Point, published in 1982 in the U.S.A., Capra comments critically, from the viewpoint of the latest findings of subatomic physics, on the mechanical-analytical ap- proach which still predominates in the biological sciences, and adds some philosophical reflections. The following quotations are important ecolo- gically and may be of interest to biologists in general: "It is now becoming apparent that overemphasis on the scientific method and on rational, analytic thinking has led to attitudes that are profoundly anti-ecological. In truth, the understanding of ecosystems is hindered by the very nature of the rational mind. Rational thinking is linear, whereas ecological awareness arises from an intuition of non-linear systems ...The Cartesian view of the universe as a mechanical system provided a 'scientific' sanction for the manipulation and exploitation of nature that has become typical of Western culture ...The problem is that scientists, encouraged by their success in treating living organisms as machines, tend to believe that they are nothing but machines ...Modern physics can show the other sciences that scientific thinking does not necessarily have to be reductionist and mechanistic, that holistic and ecological views are also scientifically sound ...

In Volume 1 of this four-volume series, ecological problems of a general nature were discussed from a global point of view. Familiarity with this is essential for a full understanding of the more specialized treatment in this and subsequent volumes, for no similar approach is to be found in other ecological handbooks for beginners. This present volume deals in detail with the special ecological relation­ ships of the tropical and subtropical zonobiomes I to III. Most ecologists proceed from the basis of their experience in the temperate zones of the northern hemisphere. As a result, many ecological writings show a certain one-sidedness and there is a danger that generalizations made will not be broadly applicable. To avoid this, particular emphasis is laid, in this vol­ urne, on the special ecological features and the characteristics of the trop­ ical and subtropical regions. More specifically, we deal not only with the relationship of the euclimatope to zonal soils and zonal vegetation, but also pay attention to azonal conditions shown in pedobiomes and in the altitudinal belts of mountains, the orobiomes. In this and the subsequent volumes the same simple scheme is followed in treating each zonobiome: 1. climate; 2. soils; 3. producers; 4. consum­ ers; 5. decomposers; 6. ecosystems; 7. sub division into biomes; 8. oro­ biomes; 9. pedobiomes and 10. zonoecotones. Where it has appeared expedient, however, we have occasionally deviated from this scheme (see Deserts D, F, G and H).

In Volume 1 of this four-volume series, ecological problems of a general nature were discussed from a global point of view. Familiarity with this is essential for a full understanding of the more specialized treatment in this and subsequent volumes, for no similar approach is to be found in other ecological handbooks for beginners. This present volume deals in detail with the special ecological relation­ ships of the tropical and subtropical zonobiomes I to III. Most ecologists proceed from the basis of their experience in the temperate zones of the northern hemisphere. As a result, many ecological writings show a certain one-sidedness and there is a danger that generalizations made will not be broadly applicable. To avoid this, particular emphasis is laid, in this vol­ urne, on the special ecological features and the characteristics of the trop­ ical and subtropical regions. More specifically, we deal not only with the relationship of the euclimatope to zonal soils and zonal vegetation, but also pay attention to azonal conditions shown in pedobiomes and in the altitudinal belts of mountains, the orobiomes. In this and the subsequent volumes the same simple scheme is followed in treating each zonobiome: 1. climate; 2. soils; 3. producers; 4. consum­ ers; 5. decomposers; 6. ecosystems; 7. sub division into biomes; 8. oro­ biomes; 9. pedobiomes and 10. zonoecotones. Where it has appeared expedient, however, we have occasionally deviated from this scheme (see Deserts D, F, G and H).