"...will no doubt assist biologists and ornithologists in their research." Audubon Naturalist Society

• 1. Introduction; 2. The study area. Ross Island and the Cape Bird penguin colonies; 3. The range of foods available to the skuas at Cape Bird during the breeding season; 4. The biomass of penguin eggs and chicks on the Northern Colony; 5. Factors of penguin biology that constrain or assist skua predation; 6. Description of scavenging and predatory behaviour of skuas and the defensive behaviour of penguins; 7. The diversity and intensity of skua foraging behaviour on the penguin colony; 8. The amount of food taken by the skuas from the penguin colony; 9. The costs and returns of foraging at the colony and at sea; 10. Immediate impact of the contestants on each other; 11. Appreciating the penguins; 12. Associating together. The longer term implications; 13. Synthesis; Appendices; References; Index.

"...will no doubt assist biologists and ornithologists in their research." Audubon Naturalist Society

• 1. Introduction; 2. The study area. Ross Island and the Cape Bird penguin colonies; 3. The range of foods available to the skuas at Cape Bird during the breeding season; 4. The biomass of penguin eggs and chicks on the Northern Colony; 5. Factors of penguin biology that constrain or assist skua predation; 6. Description of scavenging and predatory behaviour of skuas and the defensive behaviour of penguins; 7. The diversity and intensity of skua foraging behaviour on the penguin colony; 8. The amount of food taken by the skuas from the penguin colony; 9. The costs and returns of foraging at the colony and at sea; 10. Immediate impact of the contestants on each other; 11. Appreciating the penguins; 12. Associating together. The longer term implications; 13. Synthesis; Appendices; References; Index.

"...will no doubt assist biologists and ornithologists in their research." Audubon Naturalist Society

• 1. Introduction; 2. The study area. Ross Island and the Cape Bird penguin colonies; 3. The range of foods available to the skuas at Cape Bird during the breeding season; 4. The biomass of penguin eggs and chicks on the Northern Colony; 5. Factors of penguin biology that constrain or assist skua predation; 6. Description of scavenging and predatory behaviour of skuas and the defensive behaviour of penguins; 7. The diversity and intensity of skua foraging behaviour on the penguin colony; 8. The amount of food taken by the skuas from the penguin colony; 9. The costs and returns of foraging at the colony and at sea; 10. Immediate impact of the contestants on each other; 11. Appreciating the penguins; 12. Associating together. The longer term implications; 13. Synthesis; Appendices; References; Index.