In the last decades a remarkable renaissance has materialized in insect morphology, mainly triggered by the development of new cutting-edge technologies. This is an exciting time for biological synthesis where the mysteries and data derived from genomes can be combined with centuries of data from morphology and development. And, now, more than ever, detailed knowledge of morphology is essential to understanding the evolution of all groups of organisms. In this “age of phylogenomics” researchers rely on morphological data to support molecular findings, test complex evolutionary scenarios, and for placing fossil taxa. This textbook provides an in-depth treatment of the structures and the phylogeny of the megadiverse Hexapoda. The first part presents an up-to-date overview of general insect morphology with detailed drawings, scanning electron micrographs, and 3-D reconstructions. Also included is a chapter covering innovative morphological techniques (e.g., µ-computer tomography, 3-D modeling), brief treatments of insect development and phylogenetic methods, and a comprehensive morphological glossary. The second part is of a modern synthesis of insect systematics that includes taxon-specific morphological information for all Orders. The work is an invaluable reference for students and researchers working in all facets of biology and is a must for evolutionary biologists. A detailed understanding of morphology is essential in unraveling phylogenetic relationships and developing complex evolutionary scenarios. Increasingly researchers in phylogenomics are re/turning to morphological data to support their findings, while the development of new cutting-edge technologies has further increased interest in this growing field. This definitive handbook provides an in-depth treatment of insect morphology. The first part presents an up-to-date overview of insect morphology with detailed drawings, brilliant scanning electron micrographs and 3-D reconstructions as interactive PDFs. This is complemented by a chapter on innovative morphological techniques (e.g., µ-computer tomography, 3-D modeling) and a comprehensive morphological glossary. The second part treats the state of the art in insect systematics and includes taxon-specific morphological information for all orders. Systematics are treated formally, with for example the arguments for relationships (“apomorphies”) always listed explicitly. The work is a useful reference for students and researchers working in different fields of biology and a must for those dealing with insects from an evolutionary perspective.

Nannochoristidae (=Nannomecoptera) and Boreidae (=Neomecoptera) are traditionally assigned to the holometabolan Order Mecoptera. Both are presently in the focus of insect systematists. They differ strikingly from the "typical" mecopteran pattern, not only in their larval and adult morphology, but also in their life style and reproductive biology. Phylogenetic affinities between Boreidae ("snow fleas") and fleas were postulated mainly based on molecular data, and morphological characters related to food uptake and larval features suggest a close relationship between Nannochoristidae and true flies. Both groups display fascinating features such as preferred temperatures around 0°C, extremely slender larvae adapted to running water, and a highly unusual mating behaviour. Despite of the extremely low number of known species, there is no doubt that both groups are key taxa and in their own way highlights in the evolution of the extremely successful Holometabola. This and an impressive number of recent studies on various aspects of Nannochoristidae and Boreidae more than justifies a new volume in the series Handbook of Zoology dedicated to these highly unusual taxa.

Nannochoristidae (=Nannomecoptera) and Boreidae (=Neomecoptera) are traditionally assigned to the holometabolan Order Mecoptera. Both are presently in the focus of insect systematists. They differ strikingly from the "typical" mecopteran pattern, not only in their larval and adult morphology, but also in their life style and reproductive biology. Phylogenetic affinities between Boreidae ("snow fleas") and fleas were postulated mainly based on molecular data, and morphological characters related to food uptake and larval features suggest a close relationship between Nannochoristidae and true flies. Both groups display fascinating features such as preferred temperatures around 0°C, extremely slender larvae adapted to running water, and a highly unusual mating behaviour. Despite of the extremely low number of known species, there is no doubt that both groups are key taxa and in their own way highlights in the evolution of the extremely successful Holometabola. This and an impressive number of recent studies on various aspects of Nannochoristidae and Boreidae more than justifies a new volume in the series Handbook of Zoology dedicated to these highly unusual taxa.

In the last decades a remarkable renaissance has materialized in insect morphology, mainly triggered by the development of new cutting-edge technologies. This is an exciting time for biological synthesis where the mysteries and data derived from genomes can be combined with centuries of data from morphology and development. And, now, more than ever, detailed knowledge of morphology is essential to understanding the evolution of all groups of organisms. In this “age of phylogenomics” researchers rely on morphological data to support molecular findings, test complex evolutionary scenarios, and for placing fossil taxa. This textbook provides an in-depth treatment of the structures and the phylogeny of the megadiverse Hexapoda. The first part presents an up-to-date overview of general insect morphology with detailed drawings, scanning electron micrographs, and 3-D reconstructions. Also included is a chapter covering innovative morphological techniques (e.g., µ-computer tomography, 3-D modeling), brief treatments of insect development and phylogenetic methods, and a comprehensive morphological glossary. The second part is of a modern synthesis of insect systematics that includes taxon-specific morphological information for all Orders. The work is an invaluable reference for students and researchers working in all facets of biology and is a must for evolutionary biologists. A detailed understanding of morphology is essential in unraveling phylogenetic relationships and developing complex evolutionary scenarios. Increasingly researchers in phylogenomics are re/turning to morphological data to support their findings, while the development of new cutting-edge technologies has further increased interest in this growing field. This definitive handbook provides an in-depth treatment of insect morphology. The first part presents an up-to-date overview of insect morphology with detailed drawings, brilliant scanning electron micrographs and 3-D reconstructions as interactive PDFs. This is complemented by a chapter on innovative morphological techniques (e.g., µ-computer tomography, 3-D modeling) and a comprehensive morphological glossary. The second part treats the state of the art in insect systematics and includes taxon-specific morphological information for all orders. Systematics are treated formally, with for example the arguments for relationships (“apomorphies”) always listed explicitly. The work is a useful reference for students and researchers working in different fields of biology and a must for those dealing with insects from an evolutionary perspective.

Nannochoristidae (=Nannomecoptera) and Boreidae (=Neomecoptera) are traditionally assigned to the holometabolan Order Mecoptera. Both are presently in the focus of insect systematists. They differ strikingly from the "typical" mecopteran pattern, not only in their larval and adult morphology, but also in their life style and reproductive biology. Phylogenetic affinities between Boreidae ("snow fleas") and fleas were postulated mainly based on molecular data, and morphological characters related to food uptake and larval features suggest a close relationship between Nannochoristidae and true flies. Both groups display fascinating features such as preferred temperatures around 0°C, extremely slender larvae adapted to running water, and a highly unusual mating behaviour. Despite of the extremely low number of known species, there is no doubt that both groups are key taxa and in their own way highlights in the evolution of the extremely successful Holometabola. This and an impressive number of recent studies on various aspects of Nannochoristidae and Boreidae more than justifies a new volume in the series Handbook of Zoology dedicated to these highly unusual taxa.

Nannochoristidae (=Nannomecoptera) and Boreidae (=Neomecoptera) are traditionally assigned to the holometabolan Order Mecoptera. Both are presently in the focus of insect systematists. They differ strikingly from the "typical" mecopteran pattern, not only in their larval and adult morphology, but also in their life style and reproductive biology. Phylogenetic affinities between Boreidae ("snow fleas") and fleas were postulated mainly based on molecular data, and morphological characters related to food uptake and larval features suggest a close relationship between Nannochoristidae and true flies. Both groups display fascinating features such as preferred temperatures around 0°C, extremely slender larvae adapted to running water, and a highly unusual mating behaviour. Despite of the extremely low number of known species, there is no doubt that both groups are key taxa and in their own way highlights in the evolution of the extremely successful Holometabola. This and an impressive number of recent studies on various aspects of Nannochoristidae and Boreidae more than justifies a new volume in the series Handbook of Zoology dedicated to these highly unusual taxa.