Cambridge Studies in Ecology – serie

Cliffs are present in virtually every country on earth. The lack of scientific interest in cliffs to date is in striking contrast to the commonness of cliffs around the world and to the attraction cliffs have had for humans throughout history. Cliffs provide a unique habitat, rarely investigated from an ecological viewpoint. This book aims to destroy the impression of cliffs as geological structures devoid of life, by reviewing information about the geology, geomorphology, microclimate, flora and fauna of both sea and inland cliffs. For the first time, evidence is presented to suggest that cliffs worldwide may represent an invaluable type of ecosystem, consisting of some of the least disturbed habitats on earth and contributing more to the biodiversity of a region than their surface coverage would indicate.

Temperate-zone forests have been shaped by fire, wind and grazing over thousands of years. This book provides a major contribution to the study of their dynamics by considering three important themes: • The combined influence of wind, fire and herbivory on the successional trajectories and structural characteristics of forests • The interaction of deciduous and evergreen tree species to form mosaics which, in turn, influence the environment and disturbance regime • The significance of temporal and spatial scale with regard to the overall impact of disturbances These themes are explored via case studies from the forests in the Lake States of the USA (Minnesota, Wisconsin and Michigan) where the presence of large primary forest remnants provides a unique opportunity to study the long-term dynamics of near-boreal, pine and hardwood-hemlock forests. The comparability of these forests to forests in other temperate zones allows generalizations to be made that may apply more widely.

Glacier forelands provide a unique type of field laboratory for the study of landscape ecology. Their restricted physical size and the relatively simple ecosystems which they support allow comprehensive investigations to be carried out, but their particular attraction is the existence of terrain of increasing age with increasing distance from a retreating glacier, allowing inference of ecological change over long time periods. This book provides the first comprehensive review of the available information on the ecology of recently-deglaciated terrain and evaluates critically the methodology currently employed in such studies. The theme developed is that the ecology of these regions can only be understood fully by giving due consideration to the role and interaction of both physical and biological processes in the development of the landscape. By adopting this geoecological approach, the spatial variation and dynamics of vegetation and soils are considered in relation to other aspects of the landscape such as topographic variation, climate and geomorphic processes. A geoecological model is thus outlined which provides both a framework for interpreting the varied ecological nature of glacier forelands found throughout the world, and also an agenda for future research.

This book was first published in 1995, a time when, like today, there were mounting concerns that the management of our natural global heritage was failing to arrest the rapid loss of biodiversity. The problem may be traced to the extinction of enormous numbers of 'specialised' species, especially in the tropics. Tropical species either have highly particular resource requirements, such as food or nesting hollows, or specific needs for certain types of habitat, such as wetlands, making them vulnerable to changing global conditions. This book is about specialisation and generalisation in the use of resources and habitats. The author uses a broad ecological perspective to address three main questions. How do ecologists study variation in resource and habitat use, and what do we know from these studies? How well does theory account for observations, and what are the common threads between disciplines? Finally, what is the relationship between resource and habitat use? This book provides a comprehensive analysis of ecological versatility.

The adaptation of herbivore behaviour to seasonal and locational variations in vegetation quantity and quality is inadequately modelled by conventional methods. Norman Owen-Smith innovatively links the principles of adaptive behaviour to their consequences for population dynamics and community ecology, through the application of a metaphysiological modelling approach. The main focus is on large mammalian herbivores occupying seasonally variable environments such as those characterised by African savannahs, but applications to temperate zone ungulates are also included. Issues of habitat suitability, species coexistence, and population stability or instability are similarly investigated. The modelling approach accommodates various sources of environmental variability, in space and time, in a simple conceptual way and has the potential to be applied to other consumer-resource systems. This text highlights the crucial importance of adaptive consumer responses to environmental variability and is aimed particularly at academic researchers and graduate students in the field of ecology.

Mutualistic interactions between ants and plants involve rewards offered by plants and services performed by ants in a mutually advantageous relationship. The rewards are principally food and/or nest sites, and ants in turn perform a number of services for plants: they disperse and plant seeds; they protect foliage, buds, and reproductive structures from enemies such as herbivores and seed predators; they fertilize plants with essential nutrients; and they may sometimes function as pollinators. In this book, initially published in 1985, Professor Beattie reviews the fascinating natural history of ant-plant interactions, discusses the scientific evidence for the mutualistic nature of these relationships, and reaches some conclusions about the ecological and evolutionary processes that mold them. This important work explores the natural history, experimental approach, and integration with contemporary evolutionary and ecological literature of the time will appeal to a wide variety of biologists.

The Background of Ecology is a critical and up-to-date review of the origins and development of ecology, with emphasis on the major concepts and theories shared in the ecological traditions of plant and animal ecology, limnology, and oceanography. The work traces developments in each of these somewhat isolated areas and identifies, where possible, parallels or convergences among them. Dr McIntosh describes how ecology emerged as a science in the context of nineteenth-century natural histor

Mutualistic interactions between ants and plants involve rewards offered by plants and services performed by ants in a mutually advantageous relationship. The rewards are principally food and/or nest sites, and ants in turn perform a number of services for plants: they disperse and plant seeds; they protect foliage, buds, and reproductive structures from enemies such as herbivores and seed predators; they fertilize plants with essential nutrients; and they may sometimes function as pollinators. In this book, initially published in 1985, Professor Beattie reviews the fascinating natural history of ant-plant interactions, discusses the scientific evidence for the mutualistic nature of these relationships, and reaches some conclusions about the ecological and evolutionary processes that mold them. This important work explores the natural history, experimental approach, and integration with contemporary evolutionary and ecological literature of the time will appeal to a wide variety of biologists.

The rapid expansion od industry and the excessive demands made on limited natural resources have caused genuine concern at all levels of society. In the past this concern has concentrated on plants and animals and their relationships with their environments, but now attention is also turning towards microorganisms whose role is crucial to so many natural processes - from global life and mineral cycles through to the production of beer and milk products. After a brief introduction to microbiology this book concentrates on the ecological aspects of microbial life covering a wide variety of topics including structure, behaviour, growth, dispersal, interactions and how microbes act as symbionts and pathogens. Such a wide-ranging interdisciplinary approach will appeal to undergraduate and graduate students of microbiology, plant and animal ecology, agronomy, forestry and environmental sciences. Professionals working in the same fields will also find it informative as will those working in plant pathology and soil, aquatic, medical and food microbiology.

The central thesis for plant ecology is that climate exerts the dominant control on the distribution of the major vegetation types of the world. This book sets out to examine this often neglected area in two ways. In the first part the author analyses the distribution of species in relation to climate over different scales of time and place. In the second, he reviews the various approaches to explaining observed correlation between plant distribution and climate, and to establishing the mechanisms of control in physiological and biochemical terms. Dr Woodward is an authority on plant-climate relationships, and has made some of the most important contributions to our understanding of the subject. This book builds on his extensive knowledge of the subject to provide a modern text on this basic problem of plant ecology, encompassing the techniques of palaeoecology, climatology, ecology, physiology, biochemistry and genetics.

This book deals with the lifestyles of phytoplankton, the microscopic plant life living in the open waters of lakes, ponds and rivers and makes frequent references to the phytoplankton of the sea. It reviews adaptations required of organisms to live their lives independently of solid surfaces and the different ways that these have been achieved. Chapters dealing with particular topics are arranged in approximately the order in which they become a problem to the organism: How can it stay up in the water? Can it gain sufficient light and nutrients? How fast can it grow under different conditions? And what are the results of sinking out, feeding by animals, attacks by parasitic fungi and bacteria? The final chapter assembles this information to show ways in which plankton changes with season and through much longer periods of time, contrasting the behaviours in different types of lakes. This book is important for two reasons: it is the first major book on the subject for some years and projects many recent research findings; and the approach departs from previous volumes in concentrating on needs and adaptive responses to them.

Ecologists are making increasing use of computer methods in analyzing ecological data on plant and animal communities. Ecological problems naturally involve numerous variables and numerous individuals or samples. Multivariate techniques permit the summary of large, complex sets of data and provide the means to tackle many problems that cannot be investigated experimentally because of practical restraints. Ecologists are thus enabled to group similar species and similar sample sites together, and to generate hypotheses about environmental and historical factors that affect the communities. This timely book presents a full critical description of three methodologies - direct gradient analysis, ordination, and classification - from both theoretical and practical viewpoints. Both traditional and new methods are presented. Using a wide range of illustrative examples, Hugh Gauch provides an up-to-date synthesis of this field, which will be of interest to advanced students and ecologists. These mathematical tools are also used in a wide variety of other areas, from natural resource management and agronomy to the social and political sciences.

This book was originally published in 1985. In the twenty years preceding publication very considerable advances were made in our understanding of the physiology and ecology of lichens and of the functional nature of the symbiosis between the algal and fungal partners. This book provides a critical and selective review of lichen physiological ecology. The approach is a broad one, with consideration given to all the major factors in the lichen's environment and to the important processes of nitrogen fixation, photosynthesis and respiration. The book will be of interest to students and research workers in plant ecology, plant physiology and lichenology.

Deserts provide a seemingly hostile environment in which plants can, nevertheless, survive and grow. This book, originally published in 1992, considers the ecological strategies adopted by desert succulents to overcome these problems. Much fresh data on the physiological ecology of succulent plants of southern Africa is presented - the results of more than ten years' work by the authors, both in the field and the laboratory. This information forms the basis for a better understanding of the water relations and carbon balance of succulents during short- and long-term drought and leads the authors to a definition of succulence as a tool for the temporary storage of water, essential for the conservation of physiological performance during times of drought. The authors conclude by providing a basic scheme in which the significant functional aspects of succulent life strategies are given, revealing the main trends in the biology of these fascinating plants.

It is generally recognized that larger animals eat more, live longer, have larger offspring, and so on; but it is unusual to see these commonplace observations as a basis for scientific biology. A large number of empirically based relationships describe biological rates as simple functions of body size; and other such relations predict the intrinsic rate of population growth, animal speed, animal density, territory size, prey size, physiology, and morphology. Such equations almost always exist for mammals and birds, often for other vertebrates and invertebrates, sometimes for protozoa, algae, and bacteria, and occasionally even for plants. There are too many organisms to measure all aspects of the biology of every species of population, so scientists must depend on generalizations. Body size relations represent our most extensive and powerful assemblage of generalizations, but they have never been organized for use in ecology. This book represents the largest single compilation of interspecific size relations, and instructs the reader on the use of these relationships; their comparison, combination, and criticism. Both strengths and weaknesses of our current knowledge are discussed in order to indicate the many possible directions for further research. This important volume will therefore provide a point of departure toward a new applied ecology, giving quantitative solutions to real questions. It will interest advanced students of ecology and comparative physiology as well as professional biologists.

Many people learn the rudiments of the nitrogen cycle while at school, but the details of the various processes in the cycle are still not widely understood. Some of them are of great current interest, such as the use of nitrogen fixing crops to feed an increasing world population and the problems of pollution of groundwaters by nitrates. Denitrification (forming nitrogen gas) as a way of getting rid of waste from human and intensively farmed animals. This book describes the general processes of the nitrogen cycle, then gives examples of how the cycle is modified under particular ecological and geographical conditions. These examples are drawn from all the major areas of the world, and the impact of man via agriculture, forestry and fuel combustion is discussed. Numerous references are included. The book provides a background for all those whose specialist interests interact with nitrogen cycling, whether they are involved in research or have responsibility for managing the environment.

As experimental organisms, spiders offer ecologists a unique opportunity to examine the concept of the ecological community and the role which field experimentation can play in evaluating theories of population and community ecology. In this book, David Wise provides a balanced critique of field experiments designed to uncover details of spider ecology, with the dual aim of clarifying the ecology of these fascinating organisms and providing insight into the advantages and challenges of performing field experiments with a predator ubiquitous in terrestrial ecosystems. This book will be an essential reference for all ecologists wishing to learn more about the ecology of a major terrestrial predator and the use of field experimentation as a powerful technique to test ecological hypotheses.

This informative book, first published in 1987, presents the theories of community ecology within the context of a natural example. The text describes and examines issues in community ecology and shows how research on salamanders has helped to solve some of the problems surrounding the theories. Salamanders exist in stable populations of the kind assumed in community theory and are more appropriate than most other animals for research on the applications of that theory. The interesting and meaningful results, collected from observation on these excellent subjects posed challenges to beliefs within community ecology. Life histories of salamanders, fieldwork in distinctly differing habitats, competition, predation and evolution are discussed in an easily readable text. Professional ecologists and students of community ecology and herpetology will be interedted in the information synthesised in this book.

Wildfires kill many animals, but are populations of animals affected? How do animals survive the passage of fire? Why do some tree species survive and others die in a fire? Do frequent fires cause changes in plant community composition? Answering questions such as these requires an understanding of the ecological effects of fire. Aimed at senior undergraduate students, researchers, foresters and other land managers, Dr Whelan's book examines the changes wrought by fires with reference to general ecological theory. The impacts of fires on individual organisms, populations and communities are examined separately, and emphasis is placed on the importance of fire regime. Each chapter includes a listing of 'outstanding questions' that identify gaps in current knowledge. The book finishes by summarising the major aspects of ecology that are of particular relevance to management of fires - both protection against wildfires and deliberate use of fire.

A great many terrestrial plants live in close association with fungi. The features of this association, which is known as mycorrhiza, are those of a mutualistic symbiosis. Almost all plants from mycorrhizae whereby the fungus provides soil resources to the plant in exchange for energy provided by the plant. The symbiosis means greater productivity under stress for the plant and a steady energy supply for the fungus. This book addresses the diverse and complex ways in which mycorrhizae affect the mechanism for plant survival as individuals and populations, for community structure and functioning. An evolutionary/ecological approach is used to describe how and under what conditions mycorrhizal symbioses range from managing natural and agricultural lands to biotechnological processes that enhance agricultural productivity and sustainability. The Ecology of Mycorrhizae will be an invaluable book, applicable to all levels of theoretical and applied research in agronomy, botany, ecology, environmental microbiology, and plant pathology.

The theme of this book is the invasion of land by animal lines which originated in aquatic environments. It brings together physiological and ecological evidence to show both the likely routes taken out of the sea by the aquatic ancestors of terrestrial animals and the changes in structure and function associated with these routes. The author takes an ecophysiological approach, and by using representative examples, provides a novel background against which both the terrestrial adaptations of individual species and the make up and function of terrestrial ecosystems can be considered. Dr Little is the author of the highly acclaimed book The Colonisation of Land, which discusses the phylogeny and physiology of terrestrial and semi-terrestrial animals. The Terrestrial Invasion takes a fresh approach and provides an excellent introduction to the origins of land animals suitable for ecologists, physiologists and evolutionary biologists.

Wildfires kill many animals, but are populations of animals affected? How do animals survive the passage of fire? Why do some tree species survive and others die in a fire? Do frequent fires cause changes in plant community composition? Answering questions such as these requires an understanding of the ecological effects of fire. Aimed at senior undergraduate students, researchers, foresters and other land managers, Dr Whelan's book examines the changes wrought by fires with reference to general ecological theory. The impacts of fires on individual organisms, populations and communities are examined separately, and emphasis is placed on the importance of fire regime. Each chapter includes a listing of 'outstanding questions' that identify gaps in current knowledge. The book finishes by summarising the major aspects of ecology that are of particular relevance to management of fires - both protection against wildfires and deliberate use of fire.

It is almost dogma that the boreal forest in North America is a fire-dependent forest, yet ecologists often do not consider in any technical detail how forest fires produce effects on individual plants and on plant populations. Consequently, the causal connection between the behaviour of fire and its ecological consequences is poorly understood. This book sets out to correct this deficiency by assembling the relevant studies of fire intensity, rate of spread, fuel consumption, fire frequency and fire weather in the North American boreal forest. The central thesis is that the North American boreal forest has at least four wildfire characteristics that are important in understanding the dynamics of its plant populations: the large size of the burns with respect to dispersal distances, the short recurrence time of fire with respect to tree lifespans, the high mortality of plants due to the predominance of crown fires, and a good germination surface due to the large area of the forest floor which is covered by ash.

Ecological Experiments stresses the importance to ecology of field experiments, where variables are manipulated in order to collect data on specific hypotheses, as opposed to the more passive observational method. The book begins by introducing a series of ecological questions that can be addressed experimentally for example, what is the significance of competition among species? The minimal requirements of experimental design that must be met are then introduced, together with examples of good and poor experiments from the ecological literature and a consideration of the trade-offs that may be forced on the experimenter by field conditions. All ecologists, and especially students beginning their careers in field study, will find in this text a good introduction to the experimental foundation of ecology.

It is almost dogma that the boreal forest in North America is a fire-dependent forest, yet ecologists often do not consider in any technical detail how forest fires produce effects on individual plants and on plant populations. Consequently, the causal connection between the behaviour of fire and its ecological consequences is poorly understood. This book sets out to correct this deficiency by assembling the relevant studies of fire intensity, rate of spread, fuel consumption, fire frequency and fire weather in the North American boreal forest. The central thesis is that the North American boreal forest has at least four wildfire characteristics that are important in understanding the dynamics of its plant populations: the large size of the burns with respect to dispersal distances, the short recurrence time of fire with respect to tree lifespans, the high mortality of plants due to the predominance of crown fires, and a good germination surface due to the large area of the forest floor which is covered by ash.

Glacier forelands provide a unique type of field laboratory for the study of landscape ecology. Their restricted physical size and the relatively simple ecosystems which they support allow comprehensive investigations to be carried out, but their particular attraction is the existence of terrain of increasing age with increasing distance from a retreating glacier, allowing inference of ecological change over long time periods. This book provides the first comprehensive review of the available information on the ecology of recently-deglaciated terrain and evaluates critically the methodology currently employed in such studies. The theme developed is that the ecology of these regions can only be understood fully by giving due consideration to the role and interaction of both physical and biological processes in the development of the landscape. By adopting this geoecological approach, the spatial variation and dynamics of vegetation and soils are considered in relation to other aspects of the landscape such as topographic variation, climate and geomorphic processes. A geoecological model is thus outlined which provides both a framework for interpreting the varied ecological nature of glacier forelands found throughout the world, and also an agenda for future research.

Riparian Landscapes examines the ecological systems of streamside and floodplain areas from the perspective of landscape ecology. The specific spatial pattern of riparian vegetation is seen as a result of, and a control on, the ecological, geomorphological, and hydrological processes that operate along rivers. Riparian structures are controlled by the spatial dynamics of channels, flooding and soil moisture. These dynamics are part of integrated cascades of water, sediment, nutrients and carbon, to which animal and plant species respond in ways that illuminate community structure and competition. The role of the riparian zone in controlling species distribution and abundance is discussed. Intelligent management of these valuable ecological resources is highlighted. The potential for linking hydrological, geomorphological and ecological simulation models is also explored.

The mechanisms of macroevolutionary change have long been a contentious issue. Palaeoecological evidence, presented in this book, shows that evolutionary processes visible in ecological time do not build up into macroevolutionary trends, contrary to Darwin's original thesis. The author discusses how climatic oscillations on ice-age time-scales are paced by variations in the Earth's orbit, and have thus been a permanent feature of Earth history. There is, however, little evidence for macroevolutionary change in response to these climatic changes, suggesting that over geological time macroevolution does not occur as a result of accumulated short term processes. These conclusions are used to construct a post-modern evolutionary synthesis in which evolution and ecology play an equal role. Written by a leading palaeoecologist, this book will be of interest to researchers in both ecology and evolutionary biology.

The mechanisms of macroevolutionary change have long been a contentious issue. Palaeoecological evidence, presented in this book, shows that evolutionary processes visible in ecological time do not build up into macroevolutionary trends, contrary to Darwin's original thesis. The author discusses how climatic oscillations on ice-age time-scales are paced by variations in the Earth's orbit, and have thus been a permanent feature of Earth history. There is, however, little evidence for macroevolutionary change in response to these climatic changes, suggesting that over geological time macroevolution does not occur as a result of accumulated short term processes. These conclusions are used to construct a post-modern evolutionary synthesis in which evolution and ecology play an equal role. Written by a leading palaeoecologist, this book will be of interest to researchers in both ecology and evolutionary biology.