Vertebrate Paleobiology and Paleoanthropology – serie

This volume brings together faunal analysts working at many sites spanning the East African Pliocene. Although most chapters focus on the vertebrate fauna of particular localities, authors take a broad approach that seeks to compare paleoenvironmental and paleoecological patterns across localities and among various taxonomic groups. This volume aims to synthesize large amounts of faunal data, and to present the evolution of East African vertebrates in the context of environmental and climatic changes during the Pliocene.

Recent years have witnessed exciting and important scientific breakthroughs in the study of Neanderthals and their place in human evolution which have transformed our appreciation of this group’s paleobiology and evolution. This volume presents cutting-edge research by leading scientists re-examining the major debates in Neanderthal research with the use of innovative state-of-the art methods and exciting new theoretical approaches. Topics addressed include the re-evaluation of Neanderthal anatomy, inferred adaptations and habitual activities, developmental patterns, phylogenetic relationships, and the Neanderthal extinction; new methods include computer tomography, 3D geometric morphometrics, ancient DNA and bioenergetics. The diverse contributions offer fresh insights and advances in Neanderthal and modern human origins research.This is a Volume in The Max-Planck-Institute Subseries in Human Evolution coordinated by Jean-Jacques Hublin, Max-Planck-Institute for Evolutionary Anthropology, Department of Human Evolution, Leipzig, Germany

South Asia is home to a diverse range of prehistoric and contemporary cultures that include foragers, pastoralists, and farmers. In this book, archaeologists, biological anthropologists, geneticists and linguists are brought together in order to provide a comprehensive account of the history and evolution of human populations residing in the subcontinent. A wide range of topics and issues are addressed in this book, including hominin adaptations, behaviours, and dispersals; the origin and spread of food producing economies; and the cultural, biological and genetic relationship of foragers and settled communities. New theories, methodologies and interpretations presented in this book are bound to have a profound effect on the way in which the cultural record of South Asia is perceived and how this evolutionary history relates to events in the wider world.

S. E. BAILEY Department of Human Evolution Max Planck Institute for Evolutionary Anthropology Deutscher Platz 6 D-04103 Leipzig, Germany and Center for the Study of Human Origins, Department of Anthropology, New York University, 25 Waverly Place New York, NY 10003, USA sbailey@nyu. edu J. -J. HUBLIN Department of Human Evolution Max Planck Institute for Evolutionary Anthropology Deutscher Platz 6 D-04103 Leipzig, Germany hublin@eva. mpg. de When faced with choosing a topic to as teeth represent, by far, the most abundant be the focus of the first symposium material documenting different species of in Human Evolution at the Max Planck extinct non-human primates and hominins. As Institute for Evolutionary Anthropology in such, much of what we know about non- Leipzig, a paleoanthropological perspective human primate and hominin evolution is based of dental anthropology was a natural choice. on teeth. Teeth make up a disproportionate number Teeth have been a focus of interest for of the fossils discovered. They represent physical anthropologists over many gen- strongly mineralized organs of compact shape, ations. Teeth provide a multitude of which allow better preservation in geological information about humans – including deposits and archaeological sites than any cultural treatment, pathology, morphological other part of the skeleton. As a result, variation, and development. The presence of since the discoveries of the first fossils of culturally induced wear (toothpick grooves, extinct species, vertebrate paleontology has for example) reveals something about what been built primarily on analyses of teeth.

Plio-Pleistocene sites are a rare occurrence in same sites. This combination of factors is the archaeological record. When they are unique in East African Plio-Pleistocene uncovered, the faunal materials so crucial to archaeology and has stimulated much debate unlocking their behavioral meaning are often over the socioeconomic function of early sites. poorly preserved. For example, at Koobi Fora, Influential models of early hominid behavior Kenya, a prolific region that preserves several in the late 1960s and early 1970s were based classic Plio-Pleistocene sites, many bones are exclusively on information from Olduvai affected by poor cortical surface preservation Gorge (Leakey, 1971). Although Isaac’s (e. g. , (Isaac, 1997). Such taphonomic vagaries limit 1978) work at Koobi Fora expanded their the range of questions that can be addressed application, the subsequent critiques, modi- with these assemblages. In other instances, cations, and reformulations of these models access to materials can be limited due to local were based almost solely on studies from politics or rivalries between individual Olduvai (Binford, 1981, 1984; Bunn, 1981; research teams. As a result, many important Potts, 1982, 1988; Bunn and Kroll, 1986; assemblages either remain unstudied or have Blumenschine, 1995; Rose and Marshall, been interpreted without the advantage of a 1996; Domínguez-Rodrigo, 1997a, 2002; fully developed taphonomic framework, a sit- Plummer, 2004). Having said that, it is also uation that all but guarantees stagnant inter- clear that continued work at Koobi Fora and pretations.

This volume acknowledges and celebrates the contributions of Dr. Frederick S. Szalay to the field of Mammalian Evolutionary Morphology. Professor Szalay has published about 200 articles, four monographs, and six books on this subject. Throughout his career Professor Szalay has been a strong advocate for biologically and evolutionarily meaningful character analysis.  In his view, this can be accomplished only through an integrated strategy of functional, adaptational, and historical analysis. Dr. Szalay worked on several different mammalian groups during his career, and the contributions to this volume reflect his broad perspective. Chapters focus on Primates, a group to which Professor Szalay dedicated much of his career.  However, other mammalian groups on which he conducted a significant amount of research, such as marsupials and xenarthrans, are also covered in the volume.This book will be of interest to professionals and graduate students in a wide variety of related fields, including functional morphology, systematics, vertebrate paleontology, mammalogy, primatology, biological anthropology, and evolutionary biology.

The volume contains summaries of facts, theories, and unsolved problems pertaining to the unexplained extinction of dozens of genera of mostly large terrestrial mammals, which occurred ca. 13,000 calendar years ago in North America and about 1,000 years later in South America. Another equally mysterious wave of extinctions affected large Caribbean islands around 5,000 years ago. The coupling of these extinctions with the earliest appearance of human beings has led to the suggestion that foraging humans are to blame, although major climatic shifts were also taking place in the Americas during some of the extinctions. The last published volume with similar (but not identical) themes -- Extinctions in Near Time -- appeared in 1999; since then a great deal of innovative, exciting new research has been done but has not yet been compiled and summarized. Different chapters in this volume provide in-depth resumés of the chronology of the extinctions in North and South America, the possible insights into animal ecology provided by studies of stable isotopes and anatomical/physiological characteristics such as growth increments in mammoth and mastodont tusks, the clues from taphonomic research about large-mammal biology, the applications of dating methods to the extinctions debate, and archeological controversies concerning human hunting of large mammals.

An understanding of the uniquely human behavior of stone tool making tackles questions about hominins’ ability to culturally transmit and expand their base of social and practical knowledge and their cognitive capacities for advanced planning. The appearance of stone tools has often been viewed as a threshold event, impacting directly and profoundly the later course of cultural and social evolution. Alternatively, it has been understood as a prelude to significant succeeding changes in behavioral, social and biological evolution of hominins. This book presents a series of recent enquiries into the technological and adaptive significance of Oldowan stone tools. While anchored in a long research tradition, these studies rely on recent discoveries and innovative analyses of the archaeological record of ca. 2.6–1.0 million years ago in Africa and Eurasia, dealing with the earliest lithic industries as manifestations of hominin adaptations and as expressions of hominin cognitive abilities.

Michael P. Richards and Jean-Jacques Hublin The study of hominin diets, and especially how they have (primates, modern humans), (2) faunal and plant studies, (3) evolved throughout time, has long been a core research archaeology and paleoanthropology, and (4) isotopic studies. area in archaeology and paleoanthropology, but it is also This volume therefore presents research articles by most of becoming an important research area in other fields such as these participants that are mainly based on their presentations primatology, nutrition science, and evolutionary medicine. at the symposium. As can hopefully be seen in the volume, Although this is a fundamental research topic, much of the these papers provide important reviews of the current research research continues to be undertaken by specialists and there in these areas, as well as often present new research on dietary is, with some notable exceptions (e. g. , Stanford and Bunn, evolution. 2001; Ungar and Teaford, 2002; Ungar, 2007) relatively lit- In the section on modern studies Hohmann provides a tle interaction with other researchers in other fields. This is review of the diets of non-human primates, including an unfortunate, as recently it has appeared that different lines interesting discussion of the role of food-sharing amongst of evidence are causing similar conclusions about the major these primates. Snodgrass, Leonard, and Roberston provide issues of hominid dietary evolution (i. e.

There are some issues in human paleontology that seem to be timeless. Most deal with the origin and early evolution of our own genus – something about which we should care. Some of these issues pertain to taxonomy and systematics. How many species of Homo were there in the Pliocene and Pleistocene? How do we identify the earliest members the genus Homo? If there is more than one Plio-Pleistocene species, how do they relate to one another, and where and when did they evolve? Other issues relate to questions about body size, proportions and the functional adaptations of the locomotor skeleton. When did the human postcranial “Bauplan” evolve, and for what reasons? What behaviors (and what behavioral limitations) can be inferred from the postcranial bones that have been attributed to Homo habilis and Homo erectus? Still other issues relate to growth, development and life history strategies, and the biological and archeological evidence for diet and behavior in early Homo. It is often argued that dietary change played an important role in the origin and early evolution of our genus, with stone tools opening up scavenging and hunting opportunities that would have added meat protein to the diet of Homo. Still other issues relate to the environmental and climatic context in which this genus evolved.

This manuscript is the 4th Volume of the Gesher Benot Ya‘aqov (GBY) monograph sub-series. The goal of the book is to publish the lithic assemblages originating in the excavations of the Acheulian site at Gesher Benot Ya‘aqov (GBY), Israel. The authors provide the readers with detailed descriptions of the lithic assemblages, illustrations (maps, photographs, drawings) and complete inventory of the artifacts that were excavated during the seven field seasons of 1989-1997 (two in 1997) under the directorship of Prof. Naama Goren-Inbar. This manuscript includes the classification of Large Cutting Tools (bifaces: handaxes and cleavers), Cores and Core Tools, and Flake Tools made of three different raw materials (flint, basalt and limestone). This major classification system enables, in addition to the characterization of the assemblages, intra and inter assemblage analyses and comparisons. It forms the foundation and means with which the GBY cultural sequence can be investigated and compared with other Levantine, African and Asian Acheulian entities. From a methodological perspective the authors apply a detailed attribute analysis to all lithic items, a method that integrates morpho-technoogical and stylistic observations culminating in better understanding of the Acheulian realm as documented by the analysis. This analysis is aimed to refine and improve the understanding beyond that of types and their technology and to allow describing the reduction sequence (chaîne opératoire) of some of the major components of the lithic asemblages. The unique record of diverse data from GBY provides insight into hominin behavior (through time) along the margins of the paleo-Lake Hula, and sheds light on processes that led to the colonization of other parts of Eurasia. The book will be of interest to academics and students in all disciplines of Quaternary studies, and to archaeologists using GIS for intra-site spatial analysis.

This edited volume presents current archaeological research and data from the major early Acheulean sites in East Africa, and addresses three main areas of focus; 1) the tempo and mode of technological changes that led to the emergence of the Acheulean in East Africa; 2) new approaches to lithic collections, including lithic technology analyses; and 3) the debated coexistence of the Developed Oldowan and the early Acheulean. The chapters are the proceedings from the workshop titled “The Emergence of the Acheulean in East Africa”, held at University of Rome “La Sapienza” on September 12–13, 2013. The aim of the workshop was to bring together researchers currently working in this field in East Africa, in order to define the characteristics and the evolution of the early Acheulean. The volume was expanded with some chapters on the preceding Oldowan, on the African fauna and on paleovegetation, on the Acheulean in Asia and, eventually, on the Acheulean in Europe. The book is addressed to the scientific community, and will be of interest to researchers, graduate students, archaeologists, paleontologists, and paleoanthropologists. This volume is dedicated to the memory of Jean Chavaillon (March 25, 1925 - December 21, 2013), the leading archaeologist and Quaternary geologist who researched with unfailing enthusiasm the earliest human cultures and directed from 1965 to 1995 the French Archaeological Mission at Melka Kunture.

This volume brings together diverse contributions from leading archaeologists and paleoanthropologists, covering various spatial and temporal periods to distinguish convergent evolution from cultural transmission in order to see if we can discover ancient human populations. With a focus on lithic technology, the book analyzes ancient materials and cultures to systematically explore the theoretical and physical aspects of culture, convergence, and populations in human evolution and prehistory. The book will be of interest to academics, students and researchers in archaeology, paleoanthropology, genetics, and paleontology. The book begins by addressing early prehistory, discussing the convergent evolution of behaviors and the diverse ecological conditions driving the success of different evolutionary paths. Chapters discuss these topics and technology in the context of the Lower Paleolithic/Earlier Stone age and Middle Paleolithic/Middle Stone Age. The book then moves towards a focuson the prehistory of our species over the last 40,000 years. Topics covered include the human evolutionary and dispersal consequences of the Middle-Upper Paleolithic Transition in Western Eurasia. Readers will also learn about the cultural convergences, and divergences, that occurred during the Terminal Pleistocene and Holocene, such as the budding of human societies in the Americas. The book concludes by integrating these various perspectives and theories, and explores different methods of analysis to link technological developments and cultural convergence.

This volume brings together diverse contributions from leading archaeologists and paleoanthropologists, covering various spatial and temporal periods to distinguish convergent evolution from cultural transmission in order to see if we can discover ancient human populations. With a focus on lithic technology, the book analyzes ancient materials and cultures to systematically explore the theoretical and physical aspects of culture, convergence, and populations in human evolution and prehistory. The book will be of interest to academics, students and researchers in archaeology, paleoanthropology, genetics, and paleontology. The book begins by addressing early prehistory, discussing the convergent evolution of behaviors and the diverse ecological conditions driving the success of different evolutionary paths. Chapters discuss these topics and technology in the context of the Lower Paleolithic/Earlier Stone age and Middle Paleolithic/Middle Stone Age. The book then moves towards a focuson the prehistory of our species over the last 40,000 years. Topics covered include the human evolutionary and dispersal consequences of the Middle-Upper Paleolithic Transition in Western Eurasia. Readers will also learn about the cultural convergences, and divergences, that occurred during the Terminal Pleistocene and Holocene, such as the budding of human societies in the Americas. The book concludes by integrating these various perspectives and theories, and explores different methods of analysis to link technological developments and cultural convergence.

This book recognizes and celebrates the contributions of Professor Ashok Sahni to the field of paleontology. Prof. Sahni established a School of Vertebrate Palaeontology at Panjab University, Chandigarh, India, where he trained many of today’s vertebrate paleontologists of India. The book covers topics on evolutionary patterns, macroevolutionary events, origination and radiation events, changes in physical environments & climate and their implications for biodiversity dynamics, intercontinental affinities and biogeographic connections in a plate tectonic framework. The book begins by exploring India in the age of the dinosaurs, discussing new fossil remains from the Jurassic Era, then moves through the Cretaceous and Eocene to provide a picture on faunal and floral changes in Gondwanaland in the context of plate tectonics. Furthermore, the book explores the evolutionary patterns and biotic dispersals that resulted from the northward drift of Indian plate during the Cretaceous and its collision with Asia in the Eocene. The respective chapters reveal the role of plate tectonics and climate in shaping the geographical distribution of plants and animals in Gondwana, specifically in India, as well as the post-India/Asia collision implications for biodiversity changes and biogeography in the region’s continental environments. Given its scope, the book will appeal to vertebrate paleontologists, evolutionary biologists, and paleobiogeographers.

This book recognizes and celebrates the contributions of Professor Ashok Sahni to the field of paleontology. Prof. Sahni established a School of Vertebrate Palaeontology at Panjab University, Chandigarh, India, where he trained many of today’s vertebrate paleontologists of India. The book covers topics on evolutionary patterns, macroevolutionary events, origination and radiation events, changes in physical environments & climate and their implications for biodiversity dynamics, intercontinental affinities and biogeographic connections in a plate tectonic framework. The book begins by exploring India in the age of the dinosaurs, discussing new fossil remains from the Jurassic Era, then moves through the Cretaceous and Eocene to provide a picture on faunal and floral changes in Gondwanaland in the context of plate tectonics. Furthermore, the book explores the evolutionary patterns and biotic dispersals that resulted from the northward drift of Indian plate during the Cretaceous and its collision with Asia in the Eocene. The respective chapters reveal the role of plate tectonics and climate in shaping the geographical distribution of plants and animals in Gondwana, specifically in India, as well as the post-India/Asia collision implications for biodiversity changes and biogeography in the region’s continental environments. Given its scope, the book will appeal to vertebrate paleontologists, evolutionary biologists, and paleobiogeographers.

This monograph presents the results of over 10 years of paleontological and geological survey in the Baynunah Formation of the United Arab Emirates. Exposed widely in western Abu Dhabi Emirate, the Baynunah Formation and its fossils provide the only record of terrestrial environments and evolution in the Arabian Peninsula during the late Miocene epoch (12-5 Ma).This volume describes new fossils collected since 2002, presented systematically by taxon, and including mammals, reptiles, and invertebrates, as well as fossil trackways. The discoveries are framed within the results of new geological, geochemical, and geochrononological analyses, providing an updated and synthetic view of the age, environments, and biogeographic relationships of this important fossil assemblage.

This monograph presents the results of over 10 years of paleontological and geological survey in the Baynunah Formation of the United Arab Emirates. Exposed widely in western Abu Dhabi Emirate, the Baynunah Formation and its fossils provide the only record of terrestrial environments and evolution in the Arabian Peninsula during the late Miocene epoch (12-5 Ma).This volume describes new fossils collected since 2002, presented systematically by taxon, and including mammals, reptiles, and invertebrates, as well as fossil trackways. The discoveries are framed within the results of new geological, geochemical, and geochrononological analyses, providing an updated and synthetic view of the age, environments, and biogeographic relationships of this important fossil assemblage.

This edited volume provides the historical and geological background, as well as the details pertaining to the morphology and morphometric assessments, of a singular human cranial specimen from the Late Pleistocene discovered in Hofmeyr, South Africa. The chapters are divided into 4 main sections. Section 1 discusses the discovery and historical context of the skull, while section 2 addresses its geological and geochronological contexts through dating and stable isotope analyses. Section 3 details the general morphological and morphometric analyses (description, 3-D reconstruction, morphological comparisons), and section 4 details the specific morphological analyses performed (inner ear, dentition, endocranial morphology and size). The volume will be of interest to professional and student paleoanthropologists interested in the later phases of human evolution.

This edited volume provides the historical and geological background, as well as the details pertaining to the morphology and morphometric assessments, of a singular human cranial specimen from the Late Pleistocene discovered in Hofmeyr, South Africa. The chapters are divided into 4 main sections. Section 1 discusses the discovery and historical context of the skull, while section 2 addresses its geological and geochronological contexts through dating and stable isotope analyses. Section 3 details the general morphological and morphometric analyses (description, 3-D reconstruction, morphological comparisons), and section 4 details the specific morphological analyses performed (inner ear, dentition, endocranial morphology and size). The volume will be of interest to professional and student paleoanthropologists interested in the later phases of human evolution.

This volume presents an array of different case studies which take as primary material data sourced from the NOW (‘New and Old Worlds’) database of fossil mammals. The NOW database was one of the very first large paleobiological databases, and since 1996 it has been expanded from including mainly Neogene European land mammals to cover the entire Cenozoic at a global scale. In the last two decades the number of works that are based in the use of huge databases to explore ecological and evolutionary questions has increased exponentially, and even though the importance of big data in paleobiological research has been outlined in selected chapters of general works, no volume has appeared before this one which solely focuses on the databases as a primary source in reconstructing the past. The purpose of this book is to provide an illustrative volume showing the importance of big data in paleobiological research, and presenting a broad array of unpublished examples and case studies. The book is mainly aimed to professional palaeobiologists working with Cenozoic land mammals, but the scope of the book is broad enough to fit the interest for evolutionary biologists, paleoclimatologists and paleoecologists.The volume is divided in four parts. The first part includes two chapters on the development of large paleobiological databases, providing a first-hand account on the logic and the functioning of these databases. This is a much-needed perspective which is ignored by most researchers and users of such databases and, even if centered in the NOW database, the lessons that can be learned from this part can be extended to other examples. After this introductory part, the body of the book follows and is divided into three parts: patterns in regional faunas; large scale patterns and processes; and ecological, biogeographical and evolutionary patterns of key taxa. Each chapter is written by well-known specialists in the field, withsome participation of members of the NOW advisory board. The array of selected mammal taxa ranges from carnivores, equids, ruminants and rodents to the genus Homo. The topics studied also include the diversification and radiation of major clades, large-scale paleobiogeographical patterns, the evolution of ecomorphological patterns and paleobiological problems such as evolution of body size or species longevity. In most cases the results are discussed in relation to protracted environmental or paleogeographic changes.

This volume presents an array of different case studies which take as primary material data sourced from the NOW (‘New and Old Worlds’) database of fossil mammals. The NOW database was one of the very first large paleobiological databases, and since 1996 it has been expanded from including mainly Neogene European land mammals to cover the entire Cenozoic at a global scale. In the last two decades the number of works that are based in the use of huge databases to explore ecological and evolutionary questions has increased exponentially, and even though the importance of big data in paleobiological research has been outlined in selected chapters of general works, no volume has appeared before this one which solely focuses on the databases as a primary source in reconstructing the past. The purpose of this book is to provide an illustrative volume showing the importance of big data in paleobiological research, and presenting a broad array of unpublished examples and case studies. The book is mainly aimed to professional palaeobiologists working with Cenozoic land mammals, but the scope of the book is broad enough to fit the interest for evolutionary biologists, paleoclimatologists and paleoecologists.The volume is divided in four parts. The first part includes two chapters on the development of large paleobiological databases, providing a first-hand account on the logic and the functioning of these databases. This is a much-needed perspective which is ignored by most researchers and users of such databases and, even if centered in the NOW database, the lessons that can be learned from this part can be extended to other examples. After this introductory part, the body of the book follows and is divided into three parts: patterns in regional faunas; large scale patterns and processes; and ecological, biogeographical and evolutionary patterns of key taxa. Each chapter is written by well-known specialists in the field, withsome participation of members of the NOW advisory board. The array of selected mammal taxa ranges from carnivores, equids, ruminants and rodents to the genus Homo. The topics studied also include the diversification and radiation of major clades, large-scale paleobiogeographical patterns, the evolution of ecomorphological patterns and paleobiological problems such as evolution of body size or species longevity. In most cases the results are discussed in relation to protracted environmental or paleogeographic changes.

The paleontological site of Senèze (Haute Loire, central France) was discovered in 1892 inside a volcanic crater. For over 40 years, local peasant Pierre Philis collected fossils and sold them to French and Swiss museums. The site became world-famous for its well-preserved skeletons of ungulates and carnivores, as well as rare but well-preserved remains of primates and other mammals. It is considered the reference fauna for the late Villafranchian and MNQ 18 biochronological units of European mammalian evolution, but the lack of provenance data made modern research difficult. From 2000-2006, the multidisciplinary Franco-American Senèze Research Project undertook five seasons of major fieldwork, with the goals of clarifying the age, stratigraphy and taphonomy of Senèze, as well as finding additional remains, especially of the less well-known taxa. In this volume, following a history of study and summary of the new fieldwork, four geological chapters consider field methods, stratigraphy, volcanology and dating. Combining argon-argon ages and paleomagnetic calibration, the newly recovered fossils are shown to date between 2.20 and 2.08 Ma, with concentrations ca. 2.20-2.18 and 2.10-2.08 Ma, significantly older than previously thought. Chapters on palynology, ichthyology and ornithology are followed by eight chapters on the fossil mammals. The chapter on biochronology places Senèze among other sites at the start of MNQ 18, which is estimated to end ca. 1.7 Ma. Of some 2200 specimens known from the site, over half are cervids, with bovids, rhinocerotids and equids far behind. According to data from palynology and the habitat preferences of the more common mammals, the paleoenvironment around the Senèze maar would have included forest, woodland and grassland, perhaps in a warmer and moister climate than today. Taphonomic studies revealed that bones often rested a long time under water, lacked any indication of carnivore attack and often displayed pathologies in their joints. It is likely that most of the associated skeletons were preserved undisturbed after large mammals fell into the paleolake and drowned without being able to climb out. This book responds to the long-held desire of later Cenozoic paleontologists to see a modern study of a site recognized worldwide as a biochronologic reference for the Plio-Pleistocene. Our study required renewed fieldwork using up to date techniques of topography, sedimentology, stratigraphy, geochronology and taphonomy. The systematic paleontology chapters are based on re-study of the entire body of Senèze fossils collected during more than a century of research. The volume will be of interest to paleontologists, especially those concerned with the evolution of the European fauna and with the taxa studied, as well as with paleoenvironmental reconstruction and biogeography. It will also be of value to mammalogists interested in analyses of near-modern taxa and to paleoanthropologists, archaeologists and taphonomists interested in the methods utilized and the role of Senèze as a comparative standard for a site of this age without human intervention. It will surely be an essential reference for all those who want to know more about Life in Central France Around Two Million Years Ago.

The paleontological site of Senèze (Haute Loire, central France) was discovered in 1892 inside a volcanic crater. For over 40 years, local peasant Pierre Philis collected fossils and sold them to French and Swiss museums. The site became world-famous for its well-preserved skeletons of ungulates and carnivores, as well as rare but well-preserved remains of primates and other mammals. It is considered the reference fauna for the late Villafranchian and MNQ 18 biochronological units of European mammalian evolution, but the lack of provenance data made modern research difficult. From 2000-2006, the multidisciplinary Franco-American Senèze Research Project undertook five seasons of major fieldwork, with the goals of clarifying the age, stratigraphy and taphonomy of Senèze, as well as finding additional remains, especially of the less well-known taxa. In this volume, following a history of study and summary of the new fieldwork, four geological chapters consider field methods, stratigraphy, volcanology and dating. Combining argon-argon ages and paleomagnetic calibration, the newly recovered fossils are shown to date between 2.20 and 2.08 Ma, with concentrations ca. 2.20-2.18 and 2.10-2.08 Ma, significantly older than previously thought. Chapters on palynology, ichthyology and ornithology are followed by eight chapters on the fossil mammals. The chapter on biochronology places Senèze among other sites at the start of MNQ 18, which is estimated to end ca. 1.7 Ma. Of some 2200 specimens known from the site, over half are cervids, with bovids, rhinocerotids and equids far behind. According to data from palynology and the habitat preferences of the more common mammals, the paleoenvironment around the Senèze maar would have included forest, woodland and grassland, perhaps in a warmer and moister climate than today. Taphonomic studies revealed that bones often rested a long time under water, lacked any indication of carnivore attack and often displayed pathologies in their joints. It is likely that most of the associated skeletons were preserved undisturbed after large mammals fell into the paleolake and drowned without being able to climb out. This book responds to the long-held desire of later Cenozoic paleontologists to see a modern study of a site recognized worldwide as a biochronologic reference for the Plio-Pleistocene. Our study required renewed fieldwork using up to date techniques of topography, sedimentology, stratigraphy, geochronology and taphonomy. The systematic paleontology chapters are based on re-study of the entire body of Senèze fossils collected during more than a century of research. The volume will be of interest to paleontologists, especially those concerned with the evolution of the European fauna and with the taxa studied, as well as with paleoenvironmental reconstruction and biogeography. It will also be of value to mammalogists interested in analyses of near-modern taxa and to paleoanthropologists, archaeologists and taphonomists interested in the methods utilized and the role of Senèze as a comparative standard for a site of this age without human intervention. It will surely be an essential reference for all those who want to know more about Life in Central France Around Two Million Years Ago.

This edited volume describes the geology, stratigraphy, anthropology, archaeology, dating, taphonomy, paleobotany, paleontology and paleoecology of Azokh caves (also known as Azykh or Azikh). The chapters review exhaustively the key recent research on this limestone karstic site, which is located near the village of the same name in the region of Nagorno Karabagh in the south-eastern end of the Lesser Caucasus. The site is significant due to its geographic location at an important migratory crossroad between Africa and Eurasia. These caves contain an almost complete sedimentary sequence of the transition between H.heidelbergensis and H. neanderthalensis continuing to later Pleistocene and Holocene stratified sediment. The site is also important due to the discovery of Neanderthal remains by the current research group in addition to the Middle Pleistocene hominin fossils during a previous phase of excavation work led by M. Huseinov. At the heart of this book is the matter of how this site relates to human evolution.

The aim of the book is to present original and though-provoking essays in human paleontology and prehistory, which are at the forefront of human evolutionary research, in honor of Professor Yoel Rak (a leading scholar in paleoanthropology).​ ​The volume presents a collection of original papers contributed by many of Yoel's friends and colleagues from all over the globe. Contributions from experts around the globe fall roughly into three broad categories: Reflections on some of the broad theoretical questions of evolution, and especially about human evolution; the early hominins, with special emphasis on Australopithecus afarensis and Paranthropus; and the Neanderthals, that contentious group of our closest extinct relatives. Within and across these categories, nearly every paper addresses combinations of methodological, analytical and theoretical questions that are pertinent to the whole human evolutionary time span. This book will appeal most to scholars and advanced students in paleoanthropology, human paleontology and prehistoric archaeology.

This manuscript is the 4th Volume of the Gesher Benot Ya‘aqov (GBY) monograph sub-series. The goal of the book is to publish the lithic assemblages originating in the excavations of the Acheulian site at Gesher Benot Ya‘aqov (GBY), Israel. The authors provide the readers with detailed descriptions of the lithic assemblages, illustrations (maps, photographs, drawings) and complete inventory of the artifacts that were excavated during the seven field seasons of 1989-1997 (two in 1997) under the directorship of Prof. Naama Goren-Inbar. This manuscript includes the classification of Large Cutting Tools (bifaces: handaxes and cleavers), Cores and Core Tools, and Flake Tools made of three different raw materials (flint, basalt and limestone). This major classification system enables, in addition to the characterization of the assemblages, intra and inter assemblage analyses and comparisons. It forms the foundation and means with which the GBY cultural sequence can be investigated and compared with other Levantine, African and Asian Acheulian entities. From a methodological perspective the authors apply a detailed attribute analysis to all lithic items, a method that integrates morpho-technoogical and stylistic observations culminating in better understanding of the Acheulian realm as documented by the analysis. This analysis is aimed to refine and improve the understanding beyond that of types and their technology and to allow describing the reduction sequence (chaîne opératoire) of some of the major components of the lithic asemblages. The unique record of diverse data from GBY provides insight into hominin behavior (through time) along the margins of the paleo-Lake Hula, and sheds light on processes that led to the colonization of other parts of Eurasia. The book will be of interest to academics and students in all disciplines of Quaternary studies, and to archaeologists using GIS for intra-site spatial analysis.

This edited volume presents current archaeological research and data from the major early Acheulean sites in East Africa, and addresses three main areas of focus; 1) the tempo and mode of technological changes that led to the emergence of the Acheulean in East Africa; 2) new approaches to lithic collections, including lithic technology analyses; and 3) the debated coexistence of the Developed Oldowan and the early Acheulean. The chapters are the proceedings from the workshop titled “The Emergence of the Acheulean in East Africa”, held at University of Rome “La Sapienza” on September 12–13, 2013. The aim of the workshop was to bring together researchers currently working in this field in East Africa, in order to define the characteristics and the evolution of the early Acheulean. The volume was expanded with some chapters on the preceding Oldowan, on the African fauna and on paleovegetation, on the Acheulean in Asia and, eventually, on the Acheulean in Europe. The book is addressed to the scientific community, and will be of interest to researchers, graduate students, archaeologists, paleontologists, and paleoanthropologists. This volume is dedicated to the memory of Jean Chavaillon (March 25, 1925 - December 21, 2013), the leading archaeologist and Quaternary geologist who researched with unfailing enthusiasm the earliest human cultures and directed from 1965 to 1995 the French Archaeological Mission at Melka Kunture.

This edited volume describes the geology, stratigraphy, anthropology, archaeology, dating, taphonomy, paleobotany, paleontology and paleoecology of Azokh caves (also known as Azykh or Azikh). The chapters review exhaustively the key recent research on this limestone karstic site, which is located near the village of the same name in the region of Nagorno Karabagh in the south-eastern end of the Lesser Caucasus. The site is significant due to its geographic location at an important migratory crossroad between Africa and Eurasia. These caves contain an almost complete sedimentary sequence of the transition between H.heidelbergensis and H. neanderthalensis continuing to later Pleistocene and Holocene stratified sediment. The site is also important due to the discovery of Neanderthal remains by the current research group in addition to the Middle Pleistocene hominin fossils during a previous phase of excavation work led by M. Huseinov. At the heart of this book is the matter of how this site relates to human evolution.